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The tripeptide backbone (Gly-His-Lys) coordinates a single copper ion through its histidine and amine groups, which is why GHK-Cu is grouped with the copper peptides rather than the unliganded GHK sequence
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Uncoupling agents act as lipophilic weak acids, associating with protons on the exterior of mitochondria, passing through the membrane with the bound proton, and dissociating the proton on the interior of the mitochondrion
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